Substrate Meaning

"This definition [of "tool"] is not simple, but contains several elements. The tool must not be part of the animal's body (a beak is not a tool); the user must manipulate the tool in some way for it to realise its function; and, finally, a tool cannot be attached to the substrate. This is a fairly clear definition, but does seem to produce some rather arbitrary distinctions (Hansell 1987b). The spider Dinopis, for example, makes a small web which it holds in its legs, thrusting it down on passing ants. This is a tool, but all other webs, however complex, are not since they are anchored to the substrate. The woodpecker finch […] that uses a fine stick held in the beak to extract insect prey from wood, is a tool user, but a shrike […] that impales an insect on a thorn still attached to the bush is not." — 2000, Mike Hansell, Bird Nests and Construction Behaviour, →ISBN, page 90:

"Detach/subtract [tasks involve] Severing a fixed attachment between environmental objects (or the substrate) or removing object(s) from another unattached object, so the latter is a more useful tool." — 2006, Edward A. Wasserman, Thomas R. Zentall, Comparative Cognition: Experimental Explorations of Animal Intelligence, →ISBN, page 520:

"Amphisteginids and peneroplids were among the few taxa found in the bay environments, probably due to their preferences for phytal substrates and tolerance to moderate levels of eutrophication." — 2015 December 29, “Spatial Patterns in the Distribution, Diversity and Abundance of Benthic Foraminifera around Moorea (Society Archipelago, French Polynesia)”, in PLOS ONE, →DOI:

"The melted glass being supported by the substrated sand." — 1663, Robert Boyle, “(please specify the page)”, in Some Considerations Touching the Vsefulnesse of Experimental Naturall Philosophy, […], Oxford, Oxfordshire: […] Hen[ry] Hall printer to the University, for Ric[hard] Davis, →OCLC, part I: